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Introduction Jonathan Hodge and Gregory Radick Part I. Darwin's Theorising: 1. The making of a philosophical naturalist Phillip R. Sloan 2. The notebook programmes and projects of Darwin's London years Jonathan Hodge 3. Darwin on generation, pangenesis and sexual selection Jim Endersby 4. Darwin on mind, morals and emotions Robert J. Richards 5. The arguments in the Origin of Species C. Kenneth Waters Part II. Historical Contexts: 6. Is the theory of natural selection independent of its history? Gregory Radick 7. Darwin's science and Victorian philosophy of science David L. Hull 8. Darwin and Victorian Christianity John Hedley Brooke 9. Darwin, social Darwinism and eugenics Diane B. Paul 10. From Darwin to today in evolutionary biology Jean Gayon Part III. Philosophical Themes: 11. Metaphysical and epistemological issues in modern Darwinian theory Elliott Sober 12. Darwinian concepts in the philosophy of mind Kim Sterelny 13. Darwinism in moral philosophy and social theory Alex Rosenberg 14. Belief in God in a Darwinian age Michael Ruse Part IV. Ways Forward: 15. In Darwin's wake, where am I? Daniel C. Dennett 16. Ethical expressions: why moralists scowl, frown and smile Owen Flanagan 17. Giving Darwin his due Philip Kitcher.